The Adventure of the Cohen Modal Haplotype

Nearly 50 years ago W. D. Hamilton published two papers, The genetical evolution of social behaviour I & The genetical evolution of social behaviour II, which helped revolutionize our conception of how social and genetic process might work in concert. It opened up a field of research which was highlighted in Richard Dawkins' The Selfish Gene, and helped make inclusive fitness a general idea which allows us to view specific phenomena through a powerful theoretical lens. Hamilton's original work was broad in its implications and abstract in method, but concretely utilized various eusocial insects as biological illustrations, and in particular the hymenoptera, which are characterized by haplodiploid sex-determination.